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The
two plants shown on this page illustrate how much remains to be done in
botany. Determining the exact species of a plant is dependent on
the accuracy of the botanical keys available. The primary keys for
the Four Corners area (and for large areas of the West) don't always
agree and one is sometimes left with doubts about exactly what plant one
is looking at.
Descriptions of the two plants shown on this page are in conflict on a number of plant characteristics. The descriptions of Isocoma rusbyi, for instance, vary widely from the Flora of North America to Intermountain Flora to Utah Flora to Colorado Flora. For example, FNA indicates that the leaves of Isocoma rusbyi are "not resinous"; Utah Flora indicates they are "resinous"; Intermountain Flora says "glandular-glutinous"; and Colorado Flora does not describe this aspect of the plant but calls the plant, Isocoma drummondii, a name given to a plant that, according to the other botanical texts, exists only in Texas, primarily in the very southern Gulf tip. FNA indicates that the phyllaries of Isocoma rusbyi are "sparsely or not at all gland-dotted, without resin pockets"; Utah Flora says the phyllaries have a thick... spot... [and are] resinous"; Intermountain indicates that the "involucre [is] glandular-glutinous or varnished" and is straw-colored except for the "green or greenish tip"; Colorado Flora does not mention these characteristics. FNA indicates that I. rusbyi is found in "Rocky or sandy soils, sometimes in clay, usually saline, desert shrub communities, sometimes with scattered junipers"; Intermountain says "open hillsides and dry stream-beds"; Utah Flora indicates "riparian areas"; Colorado Flora says "desert-steppe , floodplains". There are, of course, agreements on some aspects of the plants, but overall I certainly have some doubts about the identification of the plants. FNA indicates that I. pluriflora is found in Arizona, New Mexico, and Texas; I. rusbyi is found just in the Four Corners states; and, as indicated above, I. drummondii is found only in Texas. (Intermountain and Utah Flora indicate that I. rusbyi and I. drummondii are "remarkably similar" and "very similar morphologically".) The genus was named by Thomas Nuttall in 1840. The name comes from the Greek which gives us "isos" for "equal" and "coma" for "mane" referring to the pappus hairs, which you can see gleaming white below the small, bulbous, individual flowers and above the green/yellow, pointed-tip phyllaries in the second and last photographs below.
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Isocoma
pluriflora. Synonym:
Haplopappus pluriflorus.
Semi-desert. Grasslands, openings
along washes. Summer. From a distance Isocoma pluriflora and Isocoma rusbyi (shown below) appear to be some form of Rabbitbrush, but a closer look shows flowers, stems, and leaves of a different structure. The plants are only about one or two feet tall. Both Isocomas shown on this page seem to prefer intermittently wet locations, primarily desert washes and road sides, but I have also seen them in drier semi-desert woodlands. The species was first named Linosyris pluriflora by Torrey and Gray in 1842. It was renamed Isocoma pluriflora by Edward Greene. "Pluri" is Latin for "more, several", thus, "many-flowered". |
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Isocoma
pluriflora. Synonym: Haplopappus pluriflorus. Semi-desert. Grasslands, openings
along washes. Summer. |
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Isocoma
pluriflora. Synonym:
Haplopappus pluriflorus.
Semi-desert. Grasslands, openings
along washes. Summer. Leaves on plants I found in De-Na-Zin were almost all prominently toothed; very few early leaves were entire. The Flora of North America indicates that I. pluriflora is "sometimes shallowly toothed" toward the tip. The bright green leaves at left are the first leaves of early spring; they will darken with age. |
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Isocoma
rusbyi. Synonym:
Haplopappus rusbyi.
Semi-desert. Grasslands, openings
along washes, roadsides. Summer, Fall. Isocoma rusbyi is woody-based with stiff, straw-colored stems supporting masses of golden-yellow disk flowers. Leaves are hairless but they can be glistening resinous, as can the phyllaries. In late summer and early fall, roadsides can have large clusters of these plants. Henry Rusby discovered this species in Holbrook, Arizona (probably in 1883) and Edward Greene named it Isocoma rusbyi in 1906. Arthur Cronquist renamed it Haplopappus rusbyi in 1994. (More biographical information.) |
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Isocoma
rusbyi. Synonym:
Haplopappus rusbyi.
Semi-desert. Grasslands, openings
along washes, roadsides. Summer, Fall. Leaves are long and narrow, gray-green, and sessile. Stems are stiff, straw-colored, and have long longitudinal striations (visible in the field with a 10 power hand lens). |
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Isocoma
rusbyi. Synonym:
Haplopappus rusbyi.
Semi-desert. Grasslands, openings
along washes, roadsides. Summer, Fall. |
Range map © John Kartesz,
County Color Key
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Range map for Isocoma pluriflora
Range map for Isocoma rusbyi |